Juan Carlos Molinero, Ph.D.

Tel.: 0431 600 4004
Fax: 0431 600 1515
Raum: 053
E-Mail: jmolinero(at)geomar.de     

GEOMAR | Helmholtz-Zentrum für Ozeanforschung Kiel
Experimentelle Ökologie I (Nahrungsnetze)
Düsternbrooker Weg 17, Alte Botanik
24105 Kiel

Wissenschaftlicher Werdegang

  • since 2009: Scientist at GEOMAR
  • 2007-2009: Post-doctoral fellow, DFG Priority Program AQUASHIFT "The impact of climate variability in aquatic ecosystems", IFM-GEOMAR, Department of Experimental Ecology
  • 2005-2007: Post-doctoral fellow, French National Institute for Agricultural Research (INRA), Thonon les Bains
  • 2004-2005: Post-doctoral fellow, Marine Station of Wimereux, Laboratory of Oceanology and Geosciences. Climate Biodiversity in European aquatic systems
  • 2003: Doctorate by the Paris VI University, Laboratory of Oceanography of Villefranche


My research interests are broadly on the ecology of pelagic ecosystems, ranging from individuals to food web structure and ecosystem function. I employ complementary approaches, i.e. field and laboratory work, data analysis and modeling to investigate general questions in plankton ecology: (i) zooplankton life histories (i.e. trade-offs, reaction norms, demography in changing environments); (ii) extinction, recovery and invasion processes in pelagic ecosystems; (iii) synergies between climate forces and human disturbances (i.e. fishing pressure, pollution) and their consequences on plankton production and pelagic food webs. Also, I am particularly interested in across system analyses, non-linear dynamics of biological systems, disturbance ecology and general laws in ecosystem functioning.

Auszeichnungen und Stipendien

  • 2008-2010 Associated Researcher at the Sir Alistair Hardy Foundation for Ocean Sciences, SAHFOS. 
  • 2008 Publication Award Leibniz-Institut für Meereswissenschaften, IFM-GEOMAR. 
  • 2005-2007 French National Institute for Agricultural Research, INRA. 
  • 2005 Japan Society for the Promotion of Sciences (JSPS). Declined in favour of the INRA award. 
  • 2004-2005 French National Institute for Biodiversity.
  • 1998-2002 National Council for Science and Technology, CONACYT


  • 2010 Life histories of crustacean gelatinous zooplankton in changing environments. DTU Aqua, Copenhagen, Danemark
  • 2009 Climate variability and pelagic ecosystems. National Taiwan Ocean University, Keelung Taiwan
  • 2009 Ecosystem indicators in European pelagic systems under varying anthropogenic and climatic forcing. Institute for Hydrobiology and Fisheries Sciences, Hamburg, Germany
  • 2008 Overview of climate effects on aquatic ecosystems in Western Europe. Institute for Chemistry and Biology of the Marine Environment. University of Oldenburg, Germany
  • 2007 Northern Hemisphere climate impact on the pelagic ecosystem of the western Mediterranean. Institute of Oceanography and Fisheries, Split, Croatia
  • 2007 Couplings between changes in the North Atlantic climate system and Western Europe aquatic ecosystems. Leibniz-Institut für Meereswissenschaften, Kiel Germany
  • 2006 Biological indicators of environmental changes: lessons from zooplankton communities, Station Marine de Wimereux, France
  • 2006 Plankton organisms as biological indicators of the functioning in aquatic ecosystems: patterns, processes and mechanisms. Swiss Federal Institute of Aquatic Science and Technology, 22 September 2006, Zurich, Switzerland
  • 2006 Climate effects on the functioning of pelagic ecosystems. Biological indicators of global warming, top-down control and bottom-up control. Thonon les Bains, France
  • 2004 Biological responses to climate change. Mechanisms and characteristic scales: from the encounter rate in plankton organisms to regime shift in marine ecosystems. Institute of Marine Sciences and Limnology, Mexico


  • 2010 Guest Lecturer. Master Course (MC) Ecology of small and large scale processes in plankton ecosystems. University of Sciences and Technologies of Lille 1 & University of Littoral Côte d’Opale, Calais, France
  • 2009 MC Current topics in Marine Ecology (Lecture: small and large scale processes in plankton ecology). Christian-Albrechts-Universität. IFM-GEOMAR
  • 2008 Guest Lecturer. PhD Course Global warming and its consequences on evolutionary ecology. Rennes, France
  • 2008 Guest Lecturer. MC Multivarite techniques for the assessment of climate effects on plankton variability. University of Sciences and Technologies of Lille 1 & University of Littoral Côte d’Opale, Calais, France
  • 2008 MC Current topics in Marine Ecology (Lectures: Numerical Ecology, Climate effects on pelagic ecosystems). Christian-Albrechts-Universität. IFM-GEOMAR
  • 2005-2006 Guest Lecturer. University of Sciences and Technologies of Lille, France. Lectures: Zooplankton ecology
  • 2005-2006 Guest Lecturer. University of Savoie, France. Lectures: Introduction to Limnology, Ecology of Lakes, Climate effects on aquatic ecosystems
  • 1999-2003 University Paris VI, Laboratoire d’Oceanography de Villefranche. MIO seminar. Lectures: Plankton ecology and taxonomy, Introduction to data analysis


  • 39th CIESM Congress. Venise, Italy, 10-14 May, 2010.
    i) Conversi A, Fonda-Umani S, Peluso P, Molinero JC, Santojanni A, Edwards M. Hypotheses on a Mediterranean Sea regime shift in the late 1980s.
    ii) Batistic M, Garic R, JC Molinero. Changes in the non-crustacean zooplankton in the Adriatic Sea underline compound effecs of anthropogenic forces and global warming
  • ASLO Ocean Sciences Meeting 2010, 22-26 February Portland, US.
    Conversi A, Fonda-Umani S, Peluso P, Molinero JC, Santojanni A, Edwards M. The Mediterranean Sea regime shift at the end of the 1980s, and its links to other European basins
  • Climate Change Effects on Fish and Fisheries, PICES International Symposium, April 26-29, Sendai, Japan.
    Molinero JC, Casini M, Dulcic J, Grbec B, Hidalgo M, Kamburska L, Lehman A, Llope M, Mira Morovic
  • Climate Change Impacts on Estuarine and Coastal Ecosystems: a Zooplankton Perspective. 28-30 June, Boulogne sur mer, France.
    Molinero JC, K Kanevceva. Non-stationary plankton dynamics in changing coastal systems
  • International Conference on Zooplankton Ecology and Aquaculture, 6-8 October, Keelung, Taiwan. JC Molinero. Zooplankton life histories in changing environments: The case of the Northern Mediterranean Sea
  • 2nd European Large Lake Symposium 2009. Vulnerability of large lakes ecosystems, monitoring, management and measures. 10-14 August, Norrtelje, Sweden.
    i) Souissi S, JC Molinero, O Anneville. Planktonic rotifers as tiny sentinels of hemispheric wide climate oscillations and eutrophication in temperate lakes
  • 3rd GLOBEC Open Science Meeting, 22-26 June 2009, Victoria, BC, Canada.
    i) JC Molinero, Batistic M, Daly Yahia MN, et al. Climate and jellyfish outbursts in the Mediterranean Sea. A synthesis through meta-analysis.
    ii) JC Molinero, Casini M, Dulcic J, et al. Climate and pelagic ecosystems in semi-enclosed marginal seas: the case of European Shelf Seas.
    iii) JC Molinero, F Ibanez, P Nival. Assessing critical windows for the development of zooplankton populations
  • ASLO Aquatic Sciences Meeting 2009, 25-30 January 2009, Nice, France.
    i) A. Conversi, S. Fonda-Umani, T. Peluso, JC Molinero and M. Edwards. Abrupt ecosystem shift in the Mediterranean Sea.
    ii) JC Molinero, M Casini, J Dulcic, et al. Assessing climate forcing on pelagic ecosystems across European Shelf Seas: a synthesis through meta-analysis.
    iii) Casini, M., Hjelm, J., Molinero, JC, et al. Trophic cascades promote threshold-like shifts in marine ecosystem dynamics
  • Water and Climate Change Congress, 14-15 October, 2008, Antwerpen, Belgium
    S. Souissi, P. Meire, M.Tackx, A. Souissi, D. Devreker, B. Mialet, F. Azémar, JC Molinero, C. Fisson & N. Bacq. Conséquences des changements climatiques sur les peuplements biologiques et leurs habitats côtiers et estuariens : cas des estuaires de l’Escaut et de la Seine
  • ICES/CIESM Workshop to compare Zooplankton Ecology and Methodologies between the Mediterranean and the North Atlantic, 27-30 October 2008, Heraklion, Crete, Greece.
    i) Daly Yahia MN, Batistic M, Davor L, Fernandez de Puelles M, Licandro P, Malej A, Molinero JC, Siokou-Frangou I, Zervoudaki S, Daly Yahia-Kéfi O. Are the outbreaks timing of Pelagia noctiluca (Forskäl, 1771) getting more frequent in the Mediterranean basin?
    ii) Licandro P, Batistic M, Daly Yahia MN, Daly Yahia O, Malej A, Molinero JC, Souissi S. Muggiaea atlantica: an atlantic indicator of hydroclimatic changes in the Mediterranean.
    iii) Molinero JC, Daly Yahia MN, Batistic M, et al. Climate and jellyfish outbreaks in the Mediterranean Sea
  • 5th World Fisheries Congress. October 20-24, 2008. Yokohama, Japan.
    i) JC Molinero, U Sommer, M Casini, J Dulcic, et al. Climate-plankton-fish interactions across semi-enclosed European Shelf Seas. What we can learn from meta-analysis?
    ii) Anneville O, Berthon V, Glippa O, Mahjoub S, Molinero JC, Souissi S. Ontogenetic die of whitefish larvae: Insights from an experimental work.
    iii) Souissi S., Hwang J.-S., Tseng L.C., Molinero JC & Chen G.C. China Coastal Current intrusions into the north and west of Taiwan during NE monsoon evidenced by satellite images and long term tracing the key indicator species Calanus sinicus (Copepoda: Calanoida)
  • Deutschen Gesellschaft für Limnologie. 22 – 26 September, Konstanz, Germany.
    i). Molinero JC, Anneville O, Souissi S, Gerdeaux D, Sommer U. Climatically driven synchronies and phase coupling events in Western Europe pelagic ecosystems
    ii). Molinero JC, Anneville O, Souissi S, Gerdeaux D, Sommer U. Quantifying synergetic effects of global warming and anthropogenic stress on the long-term dynamics of plankton communities
  • 32nd Annual Larval Fish Conference, 4-7 August 2008 Kiel, Germany.
    J JavidPour, JC Molinero, F Sommer, U Sommer. Seasonal Habitat utilisation and life history traits in the lobate Mnemiopsis leydi in the Western Baltic Sea
  • International Symposium Effects of Climate Change on the World Oceans, May 13 – 23, 2008, Gijon, Spain.
    i) ML Fernandez de Puelles, JC Molinero, L Vicente, A Morillas, J Jansá. Zooplankton time series related to North Atlantic climate changes in waters of the Balearic Sea: a case of boundary area in the central western Mediterranean
    ii) Grbec B, M Morović, JC Molinero, et al. The influence of northern hemisphere climate patterns on the Adriatic Sea pelagic ecosystem.
    iii) M Morović, B Grbec, JC Molinero, et al. Toward a better understanding of climate forcing on decadal changes in the Adriatic Sea ecosystem
  • EGU General Assembly 2008, 13 – 18 April, Vienna, Austria.
    D. Devreker, S. Souissi, J.C. Molinero, F. Nkubito. Trade-offs of the copepod Eurytemora affinis in mega-tidal estuaries. Insights from high frequency sampling in the Seine Estuary

Arbeitsgruppen und laufende Projekte

  • SIZEMIC Working Group: Testing the Generality of Elton's Rule: Comparing Aquatic and Terrestrial Ecosystems Across Environmental Conditions. Funded by the European Science Foundation. Coordinator Dr. Julia Reiss
  • SEINE AVAL Program. Working Group: Climate effects on the Seine Estuary under global change scenarios. Funded by the French Seine-Aval Program. Coordinators: Dr Benoit Laignel and Prof. Sami Souissi
  • Jellyfish dynamics under disturbed and undisturbed ecosystems. Bilateral Project Germany-Croatia. 2010-2011, Deutscher akademischer Austauschdienst (DAAD)
  • Understanding the dynamics of invasive species under contrasting marine ecosystems. The case study of the cladoceran Penilia avirostris. Bilateral Project Germany-Greece. 2010-2011, Deutscher akademischer Austauschdienst (DAAD)
  • The effect of the global warming on marine plankton: mesocosm studies and analysis of long-term data series. Bilateral Project Germany-Argentina. 2010-2011, Deutscher akademischer Austauschdienst (DAAD)
  • Meta-analysis of climate-plankton-fish interactions across semi-enclosed European Shelf Seas. Eur-Oceans Network of excellence
  • Impact of Global Changes on the Dynamic of Gelatinous Plankton in Mediterranean Pelagic Ecosystems. Eur-Oceans Network of excellence

Reviews für wissenschaftliche Artikel

  • Aquatic Sciences, Estuarine Coastal and Shelf Sciences, Fresenius Environmental Bulletin, Geophysical Research Letters, Global Change Biology, Hydrobiologia, Journal of the Marine Biological Association, Journal of Plankton Research, Marine Biology, Marine Ecology, Marine Ecology Progress Series, Oecologia, Progress in Oceanography, Zoological Studies. 


Research Articles

  1. Casini M, Bartolino V, Molinero JC, Kornilovs G. Linking fishery, trophic interactions and climate: threshold dynamics drive herring (Clupea harengus) growth in the central Baltic Sea. Marine Ecology Progress Series (Under Review).
  2. Conversi A, Fonda-Umani S, Peluso P, Molinero JC, Santojanni A, Edwards M. The Mediterranean Sea regime shift at the end of the 1980s, and its links to other European basins. PLoS Biology (Under Review).
  3. Anneville O., Berthon V, Glippa O, Mahjoub S, Molinero JC, S Souissi. Ontogenetic dietary changes of whitefish larvae: Insights from field and experimental observations. Environmental Biology of Fish (Under Review).
  4. Anneville, O., Molinero J.C., Souissi, S. and Gerdeaux, D., 2010: Seasonal and interannual variability of cladoceran communities in two peri-alpine lakes: uncoupled response to the 2003 heat wave. J Plankton Research, 32, 6, 913-925
  5. Anneville O., Souissi S, Molinero, J.C., D Gerdeaux (2009) Influences of human activity and climate on the stock-recruitment dynamics of whitefish, Coregonus lavaretus, in Lake Geneva. Fisheries Management and Ecology, 2009, 16, 492-500.
  6. Casini M., J. Hjelm, J.C. Molinero, J. Lövgren, G. Kornilovs, M. Cardinale, V. Bartolino, A. Belgrano (2009) Trophic cascades promote threshold-like sudden shifts in the functioning of a pelagic marine ecosystem. Proceedings of the National Academy of Sciences, 106, 197-202. Doi:10.1073/pnas.0806649105.
  7. Molinero J.C., Vukanic V, Lučič D, Licandro P, Souissi S, Ibanez, F, Nival P, Calbet, A, Christou ED, Daly-Yahia N, Fernandez de Puelles ML, Mazzocchi MG, Siokou-Frangou I, Vukanic D. (2009) Mediterranean marine copepods. Basin scale trends of the calanoid Centropages typicus. Hydrobiologia, 617, pp. 41-53.
  8. Molinero J.C., Buecher, E., Lucic, D., Malej, A. and Miloslavic, M. (2009) Climate and mediteranean jellyfish: Assessing the effect of temperature regimes on jellyfish outbreak dynamics. Annales: Series historia naturalis, 19 (2), Suppl. S. 11-18.
  9. Javidpour, J., Molinero JC, Lehmann, A., Hansen, T. and Sommer, U. (2009) Annual assessment of the predation of Mnemiopsis leidyi in a new invaded environment, the Kiel Fjord (Western Baltic Sea): a matter of concern? Journal of Plankton Research, 31 (7), 729-738. doi:10.1093/plankt/fbp021.
  10. Javidpour, J., Molinero J.C., Peschutter, J. and Sommer, U. (2009) Seasonal changes and population dynamics of the ctenophore Mnemiopsis leidyi after its first year of invasion in the Kiel Fjord, Western Baltic Sea. Biological Invasions, 11, 873-882. doi:10.1007/s10530-008-9300-8.
  11. Javidpour, J., Molinero J.C. and Sommer, U. (2009) Reconsidering evidence for Mnemiopsis invastion in European waters: reply. Journal of Plankton Research. doi:10.1093/plankt/fbp106.
  12. Fernández de Puelles ML, A Morillas, A Lopez-Urrutia, JC Molinero (2009) Seasonal variability of copepod abundance in the Balearic region (Western Mediterranean) as an indicator of basin scale hydrological changes. Hydrobiologia, 617, 3-16. doi: 10.1007/s10750-008-9522-x.
  13. Devreker D, Souissi S, Molinero JC, Nkubito F. (2008) Trade-offs of the copepod Eurytemora affinis in mega-tidal estuaries. Insights from high frequency sampling in the Seine Estuary. Journal of Plankton Research, 30,1329-1342; doi:10.1093/plankt/fbn086.
  14. Mahjoub S, Anneville O, Molinero JC, Souissi S, JS Hwang (2008) Feeding mechanisms and capture success of European whitefish (Coregonus lavaretus) larvae. Knowledge and Management of Aquatic Ecosystems, 388, 05, doi: 10.1051/kmae:2008008.
  15. Casini M, Lövgren J, Hjelm J, Cardinale M, Molinero JC, Kornilovs G. (2008) Multi-level trophic cascades in a heavily exploited open marine Ecosystem. Proceedings Royal Society B. doi: 10.1098/rspb.2007.1752.
  16. Molinero JC, M. Casini, Buecher E. (2008) The influence of the Atlantic and regional climate variability on the long-term changes in gelatinous carnivore populations in the northwestern Mediterranean. Limnology and Oceanography, 53, 1456-1467.
  17. Fernandez de Puelles ML, Molinero JC. (2008) Decadal changes in hydrographic and ecological time-series in the Balearic Sea (Western Mediterranean), identifying links between climate and zooplankton. ICES Journal of Marine Sciences, 65: 311-317; doi:10.1093/icesjms/fsn017.
  18. Molinero JC, F Ibanez, E Buecher, S Souissi, S Dallot, P Nival. (2008) Climate control on the long-term changes of zooplankton communities in the Northwestern Mediterranean. Global Change Biology, 14, 11-26.
    Molinero JC, F Ibanez, S Souissi, E Bosc, P Nival (2008). Surface patterns of zooplankton spatial variability detected by high frequency sampling in the NW Mediterranean. Role of density fronts. Journal of Marine Systems, 69, 271-282.
  19. Schmmitt, F, JC Molinero, S Zongo Brisar (2008) Non-linear dynamics and intermittency in a long-term copepod time-series. Communications in Nonlinear Science and Numerical Simulations, 13, 407-415.
  20. Molinero JC, Anneville O, Souissi S, Lainé L, Gerdeaux D. (2007) Decadal changes in water temperature and ecological time-series in Lake Geneva, Europe – detecting relationships with the subtropical Atlantic climate variability. Climate Research, 34, 15-23.
    Fernandez de Puelles ML, Molinero J.C. (2007) North Atlantic climate control on plankton variability in the Balearic Sea, western Mediterranean. Geophysical Research Letters, 34, L04608, doi:10.1029/2006GL028354.
  21. Anneville O, JC Molinero, S Souissi, G Balvay, D Gerdeaux (2007) Long-term changes in the copepod community of Lake Geneva. Journal of Plankton Research, 29, i49-i59.
  22. M.G. Mazzocchi, E.D. Christou, I. Di Capua, MªLuz Fernández de Puelles, S. Fonda-Umani, JC Molinero, I. Siokou-Frangou (2007) The temporal variability of Centropages typicus in the Mediterranean Sea: From seasonal to decadal scales. Progress in Oceanography, 72, 214-232.
  23. Molinero JC, O Anneville, S Souissi, D Gerdeaux (2006). Anthropogenic and climate forcing on the long-term changes of planktonic rotifers in Lake Geneva. Journal of Plankton Research, 28, 287-296.
    Molinero JC, F Ibanez, S Souissi, M Chifflet, P Nival. (2005). Phenological changes in the Northwestern Mediterranean copepods Centropages typicus and Temora stylifera linked to climate forcing. Oecologia, 145, 640-649.
  24. Bonnet D, A Richardson, R Harris, A Hirst, G Beaugrand, M Edwards, S Ceballos, R Diekman, A Lopez-Urrutia, L Valdes, F Carlotti, JC Molinero, H Weikert, W Greve, D Lucic, A Albaina, N Daly Yahia, S Fonda Umani, A Miranda, A dos Santos, K Cook, S Robinson, ML Fernandez de Puelles (2005). An overview of Calanus helgolandicus ecology in European waters. Progress in Oceanography, 65, 1-53.
  25. Molinero JC, F Ibanez, E Buecher, S Souissi, P Nival (2005). The North Atlantic climate and the North-western Mediterranean plankton variability. Limnology and Oceanography, 50, 1213-1220.
  26. Molinero JC, P Nival (2004). Spatial distribution of the copepod Centropages typicus in Ligurian Sea (NW Mediterranean). Role of surface currents estimated by Topex-Poseidon altimetry. Comptes Rendus Biologies, 327, 1103-1111.

Other literature

  1. Batistic M, Garic R, JC Molinero (2010) Changes in the non-crustacean zooplankton in the Adriatic Sea underline compound effecs of anthropogenic forces and global warming. Proceedings 39th CIESM Congress.
  2. Conversi A, Fonda-Umani S, Peluso P, Molinero JC, Santojanni A, Edwards M, (2010) Hypotheses on a Mediterranean Sea regime shift in the late 1980s. Proceedings 39th CIESM Congress.
  3. Casini M., J. Hjelm, J.C. Molinero, J. Lövgren, G. Kornilovs, M. Cardinale, V. Bartolino, A. Belgrano (2009) Switch in ecosystem functioning triggered by trophic cascades in low diverse pelagic systems: the Baltic Sea case. Globec newsletter, 15(1):38-39.
  4. Molinero JC, U. Sommer, M. Casini, J. Dulcic, M. Edwards, B. Grbec, A. Lehmann, P. Licandro, V. Melnikov, M. Morovic, F. Sommer, A. Temnykh (2008) Climate – plankton interactions across semi – enclosed European Shelf Seas. Proceedings World Fisheries Conference.
  5. Molinero JC, M Casini, E. Buecher (2008) Connecting climate variability and the long-term changes of gelatinous carnivore populations in the northwestern Mediterranean. Globec Newsletter, 14, 7-9.
  6. Javidpour J., JC Molinero, J. Peschuter, U. Sommer (2008) Mnemiopsis leidyi, the new invader of the Baltic Sea: seasonal changes and population dynamics. Globec Newsletter, 14, 10-11.
  7. Molinero JC (2008) Large contribution of the North Atlantic climate on the long-term anomalous changes of zooplankton communiteis in the Western Mediterranean. Globec Newsletter, 14(1), 76-77.
  8. Sami Souissi, JC Molinero, Gregory Beaugrand et al. (2007) Effects of global changes on aquatic ecosystems in Western Europe: role of plankton communities. Globec Newsletter, 13, 23-25.
  9. Molinero JC, Fernandez de Puelles (2007) Climate and planktonic copepods in the Balearic Sea, Western Mediterranean. Globec Newsletter, 13, 3-4.
  10. Molinero JC, S Souissi (2006) Climatic control on the Mediterranean plankton variability. Globec Newsletter, 12, 56-58.
  11. Sami Souissi, JC Molinero, P Licandro et al. Global change and aquatic ecosystems in Western Europe – role of plankton communities (In French). Final report. French National Institute of Biodiversity. 55 pp.
  12. Molinero JC, E Buecher, F Ibanez, P Nival, S Souissi (2004). Large-scale climate control on oceanographic and ecological changes in the Ligurian Sea, NW Mediterranean. Proceedings 37th CIESM Conference, 37: 403.

Book Chapters

  1. Molinero JC et al. Climate change effects on jellyfish and zooplankton. In: Life in the Mediterranean Sea: a look at habitat changes [edit. Strambler N.]. Submitted.
  2. Schmitt FG, Huang YX, Lu ZM, Zongo SB, Molinero JC, Liu YL. Analysis of nonlinear biophysical time series in aquatic environments : scaling properties and empirical mode decomposition. In: Nonlinear Dynamics in Geosciences, edited by A. Tsonis and J. Elsner, Springer, 2007, pp. 261-290.