Dr. rer. nat. Catriona Clemmesen-Bockelmann


Research Division 3: Marine Ecology
Marine Evolutionary Ecology

Room: 5 - 1.504
Phone: +49-431 600 1349
E-Mail: cclemmesen(at)geomar.de

Helmholtz Centre for Ocean Research Kiel
Wischhofstr. 1-3
D-24148 Kiel





Complete set of publications

Selected publications:

Ory, N.C., Lehmann, A., Javidpour, J., Stöhr, R. Walls, G.L., Clemmesen, C. (2020). Factors influencing the spatial and temporal distribution of microplastics at the sea surface – A year-long monitoring case study from the urban Kiel Fjord, southwest Baltic Sea. Science of the Total environment 736 ,139493.

Costalago, D., Kisten, Y., Clemmesen, C., Strydom, N.A. (2020). Growth and nutritional condition of anchovy larvae on the West and Southeast Coasts of South Africa. Mar. Ecol. Prog. Ser. 644, 119-128.

Frommel, A.Y., Hermann, B.T., Michael, K., Lucassen, M., Clemmesen, C., Hanel, R.; Reusch, T.B.H. (2020). Differential gene expression patterns related to lipid metabolism in response to ocean acidification in larvae and juveniles of Atlantic cod. Comparative Biochemistry and Physiology, Part A, 247, 110740.

Mittermayer, F.H., Stiasny, M.H, Clemmesen, C., Bayer, T., Puvanendran, V., Chierici, M., S. Jentoft, S. & T.B.H. Reusch (2019). Transcriptome profiling reveals exposure to predicted end-of-century ocean acidification as a stealth stressor for Atlantic cod larvae.
Scientific reports, 9(1), 16908.

Stiasny, M.H., Sswat, M., Mittermayer, F.H., Falk-Petersen, I.-B., Schnell, N.K., Puvanendran, V., Mortensen. A., Reusch, T.B.H., Clemmesen, C. (2019). Divergent responses of Atlantic cod to ocean acidification and food limitation. Global Change Biology 2019, 1-11. DOI: 10.1111/gcb.14554.

Casties, I., Clemmesen, C., Briski, E. (2019). Environmental tolerance of three gammarid species with and without invasion record under current and future global warming scenarios. Diversity and Distributions 25 (4), 603-612. DOI: 10.1111/ddi.12856.

Politis, S.N., Sørensen, S.R., Mazurais, D., Servili, A., Zambonino-Infante, J.-L., Miest, J.J., Clemmesen C., Tomkiewicz J. and Butts I.A.E. (2018). Molecular Ontogeny of First-Feeding European Eel Larvae. Front. Physiol. 9:1477. doi: 10.3389/fphys.2018.0147.

Jasper, C., Huwer, B., Weiland-Bräuer, N., Clemmesen, C. (2018). First record of the non-indigenous jellyfish Blackfordia virginica (Mayer, 1910) in the Baltic Sea. Helgoland Marine Research 72: 13.

Stiasny, M.H., Mittermayer, F.H., Göttler, G., Bridges, C.R., Falk-Petersen, I-B., Puvanendran, V., Mortensen, A., Reusch, T.B.H., Clemmesen, C. (2018). Effects of parental acclimation and energy limitation in response to high CO2 exposure in Atlantic cod. Scientific Reports 8: 8348.

Bornman, E., Strydom, N., Clemmesen (2018). Appraisal of warm-temperate South African mangrove estuaries as habitats to enhance larval nutritional condition and growth of Gilchristella aestaria (Family Clupeidae) using RNA/DNA ratios. Estuaries and Coasts doi.org/10.1007/s12237-018-0375-x.

Sswat, M., Stiasny, M.H., Taucher, J., Alguero-Muniz, M., Bach, L.T.; Jutfelt, F., Riebesell, U., Clemmesen, C. (2018). Food web changes under ocean acidification promote herring larvae survival. Nature Ecology and Evolution 2, 836–840.

Sswat, M., Stiasny, M.H., Jutfelt, F., Riebesell1, U. & Clemmesen, C. (2018).Growth performance and survival of larval Atlantic herring, under the combined effects of elevated temperatures and CO2. PLoS ONE 13(1): e0191947.

Königstein, S., Dahlke, F.T., Stiasny, M.H., Storch, D., Clemmesen, C., Pörtner, H.-O. (2018). Forecasting future recruitment success for Atlantic cod in the warming and acidifying Barents Sea. Global Change Biology, doi: 10.1111/gcb.13848

Franke, A., Roth, O., De Schryver, P., Bayer, T., Garcia-Gonzalez, L., Künzel, S., Bossier, P., Miest, J.J., Clemmesen, C. (2017). Poly-β-hydroxybutyrate administration during early life: effects on performance, immunity and microbial community of European sea bass yolk-sac larvae. Scientific Reports 7: 15022.

Paulsen, M., Clemmesen, C., Hammer, C., Polte, P., Malzahn, A.M. (2016). Food-limited growth of larval Atlantic herring Clupea harengus recurrently observed in a coastal nursery area. Helgol Mar Res (2016) 70:17.

Stiasny, M.H., Mittermayer, F.H., Sswat, M., Voss, R., Jutfelt, F., Chierici, M., Puvanendran, V., Mortensen, A., Reusch, T.B.H., Clemmesen, C. (2016). Ocean Acidification Effects on Atlantic Cod Larval Survival and Recruitment to the Fished Population. PLoS One, 11(8): e0155448. doi:10.1371/journal. pone.0155448.

Peck, M.A., Baumann, H., Clemmesen, C., Herrmann, J.P., Moyano, M., Temming, A. (2015). Calibrating and comparing somatic-, nucleic acid-, and otolith-based indicators of growth and condition in young juvenile European sprat (Sprattus sprattus). Jour. Exp. Mar. Biol. Ecol 471: 217-225.

Maneja, R.H., Frommel, A.Y., Browman, H.I., Geffen, A.J., Folkvord, A., Piatkowski, U., Durif, C.M.F., Bjelland, R., Skiftesvik, A.B., Clemmesen, C. (2015). The swimming kinematics and foraging behavior of larval Atlantic herring, (Clupea harengus L.), are unaffected by elevated pCO2.Journal of Experimental Marine Biology and Ecology, 466:42-48.

Schade, F.M, Clemmesen, C., Wegner, K. M. (2014). Within- and trans- generational effects of ocean acidification on life history of marine three-spined stickleback (Gasterosteus aculeatus). Mar. Biol. DOI 10.1007/s00227-014-2450-6

Maneja, R.H., Dineshram, R., Thiyagarajan, V., Skiftesvik, A.B., Frommel, A.Y., Clemmesen, C., Geffen, A.J., Browman, H.I. (2014). The proteome of Atlantic herring (Clupea harengus L.) larvae is resistant to elevated pCO2. Mar. Poll. Bull: doi.org/10.1016/j.marpolbul.2014.07.030

Frommel, A.Y., Maneja, R.H., Lowe, D., Pascoe, C.K., Geffen, A.J., Folkvord, A., Piatkowski, U., Clemmesen, C. (2014). Organ damage in Atlantic herring larvae as a result of ocean acidification. Ecol. Appl. 24(5): 1131–1143.

Paulsen, M., Hammer, C., Malzahn, A.M, Polte, P., von Dorrien, C., Clemmesen, C. (2014). Nutritional situation for larval Atlantic herring (Clupea harengus L.) in two nursery areas in the western Baltic Sea. ICES J. Mar. Sci. 71 (4), 991-1000.

Paulsen, M., Clemmesen, C., Malzahn, A.M. (2014). Essential fatty acid (docosahexaeonic acid, DHA) availability affects growth of larval herring in the field. Marine Biology 161:239–244.

Maneja, R.H., Frommel, A.Y., Geffen, A.J., Folkvord, A., Piatkowski, U., Chang, M.Y., Clemmesen, C. (2013). Effects of ocean acidification on the calcification of otoliths of larval Atlantic cod, Gadus morhua L.. Mar. Ecol. Prog. Ser. 477: 251–258.

Voss, R., Peck, M.A., Hinrichsen, H.-H., Clemmesen, C., Baumann.H., Stepputtis, D., Bernreuter, M., Schmidt, J.O., Temming, A., Köster, F.W (2012). Recruitment processes in Baltic sprat - a re-evaluation of GLOBEC-Germany hypotheses. Progress in Oceanography 107, 51-79.

Frommel, A.Y., Maneja, R., Lowe, D., Malzahn, A., Geffen, A.J., Folkvord, A., Piatkowski, U., Reusch, T.B.H., Clemmesen, C. (2012). Severe tissue damage in Atlantic cod larvae under increasing ocean acidification. Nature Climate Change 2, 42-46.

Franke, A. and Clemmesen, C. (2011). Effect of ocean acidification on early life stages of Atlantic herring (Clupea harengus L.). Biogeosciences 8, 3697-3707.

Huwer, B., Clemmesen, C., Grønkjær, P. and Köster, F.W. (2011). Vertical distribution and growth performance of Baltic cod larvae – Field evidence for starvation-induced recruitment regulation during the larval stage? Progress in Oceanography 91, 382-396. dx.doi.org/10.1016/j.pocean.2011.04.001

Meyer, S., Caldarone, E.M., Chicharo, M.A., Clemmesen, C., Faria, A.M., Faulk, C., Folkvord, A., Holt, G.J., Hoie, H., Kanstinger, .P, Malzahn, A., Moran, D., Petereit, C., Stottrup, J.G., Peck, M.A. (2011). On the edge of death: Rates of decline and lower thresholds of biochemical condition in food-deprived fish larvae and juveniles. J. Mar. Syst. 93, 11-24. Doi: 10.1016/j.jmarsys.2011.09.010

Hinrichsen, H.H., Voss, R., Huwer, B., Clemmesen, C. (2010). Variability of larval Baltic sprat  (Sprattus sprattus L.) otolith growth: a modeling approach combining spatially and temporally resolved biotic and abiotic environmental key variables. Fish. Oceanogr. 19:6, 463-479.

Working group 'Larval ecology and aquaculture'

Scientific career

1978              Studium Diplom-Biologie, Universität Kiel
1982 - 1983   Fulbright Stipendiatin University of Washington, Seattle, USA
1983              Bachelor of Science in Fisheries, University of Washington, Seattle, USA
1985              Diplom in Biology, University of Kiel
1986              Doktorandenstipendium der CAU Kiel
1986 - 1987   Doktorandenstelle im DFG Projekt, Universität Hamburg
1988              Mutterschutz / Elternzeit
1989 - 1990   Doktorandenstelle im BMBF Projekt, Universität Hamburg
1991              Mutterschutz / Elternzeit
1992              Doktor der Naturwissenschaften in Fischereibiology, Universität Hamburg
1992 - 1994   Postdoc Position DFG Projekt, Universität Hamburg
1994 - 1996   Postdoc Position, EU Projekt Universität Kiel
1997 - 1999   Postdoc Position, EU Projekt Universität Kiel
2000 - 2002   Postdoc Position, EU Projekt MACOM, Universität Kiel
2003 - 2005   Postdoc Position, BMBF Projekt GLOBEC, Universität Kiel
2005 - 2006   DFG Forschungsstipendium für einjährigen Forschungsaufenthalt an der University
                      of Rhode Island, Narragansett, Rhode Island, USA
2007              Studie für das EU Parlament über „Climate Change and Fisheries
2008 - 2012  Teilprojektleiterin in EU Projekt EPOCA im Teilprojekt
                      Effects of ocean acidification on performance, reproduction and growth“
2008 - 2012  Teilprojektleiterin im Marie Curie Training Network CALMARO
2009 - 2012  Teilprojektleiterin im BMBF Projekt BIOACID
2009 - 2012  Koordinatorin  des WTSH Projektes NEMO
2012             50% Haushaltsstelle am GEOMAR (FB3/Evolutionsökologie mariner Fische)
2013 -           Koordinatorin  des WTSH Projektes FINEAQUA
2013 -           Teilprojektleiterin im BMBF Projekt BIOACID2
2013 -           Koordinatorin  des LLUR Projektes  OSTSEEZOOPLANKTON
2014 -           EU-BONUS Projekt BIOC3, Taskleader: Pysiological tolerance, preference
                     and phenotypic plasticity“
2014 -           Koordinatorin des LUNG Projektes Zooplankton MSRL
2014 -           POF Project „OCEANS“: „Ecological and evolutionary effects of stress regimes
                     across scales“ Workpackage Leitung zusammen mit Martin Wahl und Frank Melzner
2018-           Joint project CUSCO - "Coastal Buoyancy in a Changing Ocean.
                     Pelagic food web structures, trophic interactions and the Role of fish for trophic transfer
                     efficiency in Humboldt's Buoyancy area off Peru (WP4)".
2019-            BLE Project UFOTriNet "Development of a Trilateral Network for a automatic, continuous
                     and non-invasive  monitoring of fish stocks through the development of a
                     mobile (UFOmobil) as well as a portable underwater fish observatory (UFOportable) and
                     their  coupling with an already existing stationary underwater fish tank Observatory
                     (UFOstationary) in the Kiel Fjord".

Research focus

  • Marine Ecology
  • Experimental ecology,
  • Cultivate marine organism
  • Larval fish ecology and fisheries recruitment
  • Biological effects of global climate change
  • Ocean acidification and food webs
  • Microplastic and fish
  • Transgenerational effects and genitc research
  • Aquaculture: early life stages, probiotic bacteria, food supplements
  • Otolith organ analyses, biochemical indicators
  • Time series analyses (Kiel Fjord, Bornholm Basin, eastern Baltic Sea)

Institutional positions and functions

Head of the GEOMAR FB3 working group "Fish larval ecology and aquaculture"
Head of GEOMAR FB3 Laboratory for Biochemical Analytics
Head of the GEOMAR FB3 central repository for formol samples (biodiversity repository)
1998 - Member of the ICES Working Group on Recruitment processes
2000 - 2005 Member of the Staff Council of IFM and IFM-GEOMAR
2002 - Member of the Faculty Committee at the CAU, Kiel
2003 - Member of the German Scientific Commission for Marine Research (DWK)
2008 - Member of the Excellence Cluster "The Future Ocean", Kiel
2008 Organization and direction of the 32nd Annual Larval Fish Conference in Kiel 2010 Member of the Board of the Annette-Barthelt-Stiftung
2010 -2012 Secretary of the Early Life History Section of the American Fisheries Society
2012 -2014 President of the Early Life History Section of the American Fisheries Society
2012 - 2014 Scientific representative on the GEOMAR Scientific Council
2014 - Member of the WEB (Woman's Executive Board) Board of GEOMAR

Teaching and co-supervision of thesis

Since 1996, I represent the marine ecology with the theme "larval fish ecology" in teaching through lectures and practical exercises in the field of experimental work.
Since 1995 in collaboration with my respective superiors,  I supervised a total of 52 theses:

Müller, J. 2020: Small-scale population differentiation in Atlantic herring (Clupea harengus L.) (Masterarbeit), Christian-Albrechts-Universität Kiel, Kiel, Germany, 50 pp.
Mittermayer, F. 2018: Effects of Ocean Acidification on the transcriptome of larval Atlantic cod and impacts of parental acclimation. Dissertation, Christian-Albrechts-Universität Kiel, Kiel, Germany, pp. 144.
Stiasny, M.H. 2017: Effects of Ocean Acidification on Atlantic Cod Larvae (Gadus morhua). Dissertation, Christian-Albrechts-Universität Kiel, Kiel, Germany, pp. 102
Sswat, M. (2017) Direct and indirect effects of ocean acidification on early life stages of Atlantic cod and herring. Dissertation, Christian-Albrechts-Universität Kiel, Kiel, Germany, pp. 174
Matthias Paulsen (2017) Prey quantity and quality effects on  larval Atlantic herring (Clupea harengus L.) growth in the western Baltic Sea. Dissertation, Universität Hamburg, pp. 185
Franke A. (2017) Immunostumulation in Early Life Stages of European Sea Bass (Dicentrarchus labrax). Doktorarbeit/PhD, Christian-Albrechts-Universität Kiel, Kiel, Germany, 143 pp. 
Berghoff, L. (2016) Seasonal condition growth and performance of Eastern Baltic cod larvae based on RNA/DNA ratios (Masterarbeit), Christian-Albrechts-Universität Kiel, Kiel, Germany, 74 pp.
Yan, H. (2016) Effects of High CO2 on Immune Response and Scale Structures in Juvenile Turbot (Scophthalmus maximus) (Masterarbeit), Christian-Albrechts-Universität Kiel, Kiel, Germany, 61 pp.
Wunderow, H. (2016) The impact of ocean acidification on the skeletal ossification in herring larvae (Clupea harengus, L.) (Masterarbeit), Christian-Albrechts-Universität Kiel, Kiel, Germany, 82 pp.
Beckmann, R. (2015): Effects of microzooplankton on the planktonic food web. Master thesis, University of Kiel, 67pp.
Webers, F. (2014). Occurrence of fish larvae in the Kiel Canal and the Kiel Fjord  in relation to water temperature. Bachelor Thesis, University of Kiel, pp. 48.
Listmann, L. (2014). Gene  expression analysis in Atlantic Herring (Clupea harengus ) in response to  temperature and  CO2. Master Thesis, University of Kiel, pp. 52.
Stiasny, M. (2013). The Effect of Ocean Acidification and Warming on Survival and Growth of Atlantic Cod (Gadus morhua) Larvae. Master thesis, University of  Kiel, pp. 52.
Maneja, R. (2012). Influence of ocean acidification on otolith calcification and behaviour in fish larvae. Doktorarbeit/PhD University of Kiel, pp. 127.
Hesse, C. (2012). Analyses of growth and survival of herring larvae in the Kiel Canal. Diploma thesis, University of Kiel, pp. 62.
Müller, F. M. (2012). Einfluss von Ozeanversauerung auf die Mortalität der Embryonalstadien und den Schlupf der Larven von Gadus morhua und Pleuronectes platessa aus der westlichen Ostsee. Bachelor Arbeit Universität Kiel, pp.78.
Ecker, U. (2012): Auswirkungen von erhöhten Wintertemperaturen auf die Entwicklung des Mesozooplanktons und des Ichthyoplanktons im Frühjahr in der Kieler Förde. Bachelor Arbeit Universität Kiel, pp. 56.
Steffen, G.T. (2012): Effects of ocean acidification and paternity on the early life stages of Atlantic cod (Gadus morhua) from the south-east Canadian shelf.  Diploma thesis, University of Kiel, pp. 81.
v. Dewitz, B. (2012). A historical reconstruction of the Baltic Sea pH environment experienced by marine organisms.  Diploma thesis University of Kiel, pp. 87.
Bodenstein, S. (2012): Effects of ocean acidification and temperature on chloride cells in Atlantic herring (Clupea harengus) embryos and larvae. Diploma thesis, University of Kiel, pp. 77.
Frommel, A.: 2012: Effect of ocean acidification on the growth and development of cod larvae (Gadus morhua), Dissertation University of Kiel, pp. 107.
Form, A. (2011): Influence of anthropogenic climate change on the ecophysiology of the cold-water coral Lophelia pertusa. Dissertation, CAU, Kiel, pp. 218.
Haslob, H. (2011): Reproductive ecology of Baltic sprat and its application in stock assessment. Dissertation, Universität Kiel, pp. 133.
Paulsen, M. (2010): Einfluss der Fettsäuren auf die Kondition der Heringslarven. Diplomarbeit, Universität Kiel, pp. 112.
Grün, C. (2008): Alteration of the physiological stress tolerance of the ocean Quahog Arctica islandica during ageing processes‟, Diplomarbeit, Universität Darmstadt, pp. 114.
Petereit, C. (2009): Influence of temperature and salinity on sprat (Sprattus sprattus) eggs and yolk sac larvae from contrasting environments. Dissertation Universität Kiel, pp. 113.
Huwer, B. 2009: The recruitment process in Baltic cod. Spatio-temporal variation in starvation and predation during early life stages with special consideration of the invasive ctenophore Mnemiopsis leidyi.  Dissertation, Aarhus University, Denmark.
Gutperlet, R. (2008): Ökophysiologische Studien an der Kaltwasserkoralle Lophelia pertusa (Scleractinia). Diplomarbeit, Universität Kiel, pp. 92.
Sähn, N. (2008): Temperature effects on Baltic larval cod (Gadus morhua L.) derived from laboratory experiments. Diplomarbeit, Universität Kiel, pp.76.
Franke, A. (2008): Zum Einfluss des CO2-Partialdrucks auf die Embryonalentwicklung und den Zustand der Dottersacklarven des Atlantischen Herings, Clupea harengus. Diplomarbeit, Universität Kiel, pp. 62.
Hauss, H. (2008): Impacts of prey field characteristics on foraging and condition of larval herring (Clupea harengus). Diplomarbeit, Universität Kiel, pp. 89.
Peschutter, J. (2008): Ernährungszustand von Fischlarven in der Kieler Förde und im Nord-Ostsee-Kanal. Diplomarbeit, Universität Kiel, pp. 121.
Helland, I.P. 2008: Ecology of a sympatric pair of coregonid fish: Species interactions and temperature. Dissertation, University of Oslo, Norway.
Frommel, A. (2008): Influence of temperature on growth and biochemical-based indicators of growth in juvenile Gobiids of the Baltic Sea. Master thesis in Biological Oceanography, University of Southern Denmark, Odense in cooperation with the Leibniz Institute for Marine Sciences, Kiel, Germany, pp.82.
Stransky, J. (2007): Die Bedeutung hydrographischer Veränderungen für die zeitliche Entwicklung des Fischlarven- und Evertebratenplanktons in der Kieler Förde. Diplomarbeit, Universität Kiel, pp. 70.
Silber, Y. (2006): Einfluß von Schwermetallen (Kupfer) auf die ökophysiologische Anpassungsfähigkeit von Miesmuscheln (Mytilus spp.) verschiedener europäischer Küstengewässer an kurzfristige Salzgehaltsänderungen. Diplomarbeit, Universität Rostock, pp. 120.
Harrer, D. (2006): Experimente zum Einfluß von Temperatur auf Wachstum und Überleben von Heringslarven (Clupea harengus L.). Diplomarbeit, Universität Kiel, pp. 158.
Donner, M. (2006): Wachstum und Kondition von Heringslarven (Clupea harengus L.) in der Kieler Förde und im Nord-Ostsee- Kanal. Diplomarbeit, Universität Kiel, pp. 72.
Buehler, V. (2004): Influence of maternal effect and environmental conditions on growth and survival of Atlantic cod (Gadus morhua L.). Dissertation, Universität Kiel, pp. 139.
Holtappels, M. (2004): Nutritional condition of sprat and sardine larvae in the frontal systems of the German Bight. Diplomarbeit, Universität Rostock, pp. 86.
Huwer, B. (2004): Larval growth of Sardina pilchardus and Sprattus sprattus in relation to frontal systems in the German Bight. Diplomarbeit, Universität Kiel, pp.107.
Kamal, M.M. (2004): Growth and condition of sprat (Sprattus sprattus) larvae inferred from otolith microstructure analysis and RNA/Dna ratio in the Bornholm Basin (Central Baltic sea) during spawning season 2001. Dissertation, Universität Kiel, pp. 102.
Petereit, C. (2004): Experimente zum Temperatureinfluß auf frühe Entwicklungsstadien des Ostseedorsches Gadus morhua. Diplomarbeit, Math.-Nat. Fak., Uni. Kiel, pp. 45.
Kim, S.-K. (2003): Ecophysiological requirements for artificial seed production of Saxidomus purpuratus (Veneridae, Eulamellibranchia), a new candidate for marine aquaculture. Dissertation: Math.-Nat. Fak., Uni. Kiel, pp.157.
Röhrscheidt, H. (2001): Ernährungszustand und Wachstum von Ichthyoplankton auf der Großen Meteorbank. Diplomarbeit, Math.-Nat. Fak., Uni. Kiel, pp. 92.
Melzner, F. (2001): Determination of biochemical indices for instananous growth in juvenile cephalopods. Diplomarbeit, Math.-Nat. Fak., Uni. Kiel, pp. 90.
Malzahn, A. (2001): Untersuchungen zum Wachstum und Mortalität von Coregonen –Larven in Abhängigkeit von verschiedenen Temperaturen. Diplomarbeit, Math.-Nat. Fak., Uni. Kiel, pp. 55.
Willert, B. (2000): Kondition und Wachstum von Heringslarven in einem Mesokosmos. Diplomarbeit, Math.-Nat. Fak., Uni. Kiel, pp. 126.
Bühler, V. (1999): A study on the growth of herring larvae. Diplomarbeit,  Math.-Nat. Fak., Uni. Kiel, pp. 74.
Hoch, C. (1998): Beziehung zwischen Nukleinsäuregehalt, Wachstum und Kondition von Heringslarven. Diplomarbeit, Math.-Nat. Fak., Uni. Kiel, pp. 65.
Jung, T. (1996): Einfluß der Futterqualität auf die Entwicklung und den Ernährungszustand von Dorschlarven (Gadus morhua L.) in Aufzuchtexperimenten. Diplomarbeit, Math.-Nat. Fak., Uni. Kiel, pp. 65.
Doan, T. (1995): Wachstumsuntersuchungen an marinen Fischlarven auf der Basis von Otolithenstruktur und biochemischen Ernährungsindikatoren. Diplomarbeit, Math-Nat. Fak., Uni. Kiel, pp. 54.